Varade, Marissa L., Francis Choi, Brian Helmuth and Steven Scyphers. 2023. Catching versus counting: comparing the pro-environmental attitudes, behaviors and climate concerns of recreational fishers and citizen scientists. Sustainability 15(1), 307. doi: 10.3390/su15010307


Mancuso, Francesco Paolo, Chiara Giommi, Maria Christina Mangano, Laura Airoldi, Brian Helmuth and Gianluca Sarà. 2022. Evenness, biodiversity, and ecosystem function of intertidal communities along the Italian coast: experimental short-term response to ambient and extreme air temperatures. Science of the Total Environment, 858, 160037. 

Salois, S., T. Gouhier, B. Helmuth, S. Benson, E. Hosteler, F. Choi, N. Weller, C. Nickerson, K. Todd, and D. Cavalier. 2022. Citizen Science, civics, and resilient communities: Informing community resilience policies through local knowledge, community values, and community-generated data. Citizen Science: Theory and Practice, 7(1): 33, pp. 1-18. doi: 10.5334/cstp.516

Sittenfeld, D., Farooque, M., Helmuth, B.Benson, S., Hostetler, E., Choi, F., Weller, N., Nickerson, C., Todd, K. and Cavalier, D. 2022. Citizen Science, Civics, and Resilient Communities: Informing Community Resilience Policies Through Local Knowledge, Community Values, and Community-Generated Data. Citizen Science: Theory and Practice, 7(1): 33, pp. 1–18.

Dong, Yun-Wei, Jorge García Molinos, Eric R. Larson, Qiang Lin, Xuan Liu, Gianluca Sarà, Qing-Hua Cai, Zhixin Zhang, Brian Helmuth and Amanda Bates. 2022. Biological traits, geographic distributions, and species conservation in aquatic ecosystems. Diversity and Distributions 28: 1516-1523. doi: 10.1111/ddi.13600

Al-Mudaffar Fawzi, Nadia, Clare M. Fieseler, Brian Helmuth, Alexandra Leitão, Mehsin Al-Ainsi, Mohammad Al Mukaimi, Mohammad Al-Saidi, Fahad Al Senafi, Ivonne Bejarano, Radhouan Ben-Hamadou, Josh D’Addario, Ahmad Mujthaba Dheen Mohamed, Bruno W. Giraldes, Lyle Glowka, MAggie D. Johnson, Brett P. Lyons, Daniel Mateos-Molina, Christopher D. Marshall, Sayeed Mohammed, Pedro Range, Mohammad Reza Shokri, John M. K. Wong, Nicholas D. Pyenson. 2022. Diplomacy for the world’s hottest sea. Science 376: 1389-1390. doi: 10.1126/science.add2508. (Journal IF: 47.73)

Mangano, M.C., M. Berlino, L. Corbari, G. Milisenda, M. Lucchese, S. Terz, M. Bosch-Belmar, M.S. Azaza, J.M.F. Babarro, R. Bakiu, B.R. Broitman, A.H. Buschmann, R. Christofoletti, Y. Dong, B. Glamuzina, O. Luthman, P. Makridis, A.J.A. Nogueira, M. G. Palomo, R. Dineshram, P. SanchezJerez, H. Sevgili, M. Troell, K.Y. AbouelFadl, M. N. Azra, P. Britz, E. Carrington, I. Celíc, F. Choi, C. Qin, M.A. Dionísio, T. Dobroslavic, P. Galli, D. Giannetto, J.H. Grabowski, B. Helmuth, M.J.H. Lebata-Ramos, P.T. Lim, Y. Liu, S.M. Llorens, S. Mirto, M. Pécarevic, C. Pita, N. Ragg, E. Ravagnan, D. Saidi, K. Schultz*, M. Shaltout, S.H. Tan, V. Thiyagarajan, G. Sará. 2022. The aquaculture supply chain in the time of covid-19 pandemic: Vulnerability, resilience, solutions and priorities at the global scale. Environmental Science and Policy 127: 98-110 doi: 10.1016/j.envsci.2021.10.014. (Journal IF: 5.58).


Hassan Ali, Malik, Nadia Al-Mudaffar Fawzi, Hanaa Hussein Mohammed, Brian Helmuth, and Amanda M. Dwyer. 2021. Winners and losers: post conflict biodiversity in the stressed ecosystem of Khor Al-Zubair, Iraq. Pakistan Journal of Marine Sciences 30(2): 76-95.

Coley, John D., Nicole Betz, Brian Helmuth, Keith Ellenbogen, Steven Scyphers and Daniel Adams. 2021. Beliefs about human-nature relationships and implications for investment and stewardship surrounding land-water systems. Land, 10(12): 1293. (Journal IF: 3.40).

Montes, E., J.S. Lefcheck, E. Guerra-Castro, E. Klein, M.T. Kavanaugh, A. Azevedo Mazzuco, G. Bigatti, C.A.M.M. Cordeiro, N. Simoes, E.C. Macaya, N. Moity, E. Londoño-Cruz, B. Helmuth, F. Choi, E.H. Soto, P. Miloslavich and F.E. Muller-Karger. 2021. Optimizing large-scale Helmuth CV page 3 biodiversity sampling effort: Toward an unbalanced survey design. Oceanography. doi: 10.5670/oceanog.2021.216 (Journal IF: 2.34).

Bravo, G., N. Moity, E. Londoño-Cruz, F. Muller-Karger, G. Bigatti, E. Klein, F. Choi, L. Parmalee, B. Helmuth, and E. Montes. 2021. Robots versus humans: Automated annotation accurately quantifies essential ocean variables of rocky intertidal functional groups and habitat state. Frontiers in Marine Science 8: 691313. doi: 10.3389/fmars.2021.691313. (Journal IF: 4.44)

Cryan, A., F. Choi, D. Adams and B. Helmuth. 2021. Heat budget model facilitates exploration of thermal ecology on urban shoreline infrastructure. Ecological Engineering 171: 106371. (Journal IF: 4.04).

Muller-Karger, F., M. Kavanaugh, K. Iken, E. Montes, F. Chavez, H. Ruhl, R. Miller, J. Runge, J. Grebmeier, L. Cooper, B. Helmuth, E. Escobar-Briones, N. Hammerschlag, M. Estes, J. Pearlman, E. Hestir, E. Duffy, K.J. Sarri, C. Hudson, J. Landrum, G. Canonico, L. Jewett, J. Newton, B. Kirkpatrick, C. Anderson, A. Bates, I. Sousa-Pinto, M. Nakaoka, and J. Soares. 2021. Marine Life 2030: Forecasting changes to ocean biodiversity to inform decision making: A critical role for the Marine Biodiversity Observation Network (MBON). Marine Technology Society Journal 55(3): 84- 85. Doi: 10.4031/MTSJ.55.3.28.

Caviglia-Harris, J., K. E. Hodges, B. Helmuth, E. M. Bennett, K. Galvin, M. Krebs, K. Lips, M. Lowman, L. A. Schulte Moore and E.A.G. Schuur. 2021. The six dimensions of collective leadership that advance sustainability objectives: rethinking what it means to be an academic leader. Ecology and Society, 26(3): 9. doi 10.5751/ES-12396-260309. (Journal IF: 4.40).

Minuti, J.J., C.A. Corra, B.S. Helmuth, and B.D. Russell. 2021. Increased thermal sensitivity of a tropical marine gastropod under combined CO2 and temperature stress. Frontiers in Marine Science, 8:643377. Doi: 10.3389/fmars.2021.643377. (Journal IF: 4.44).

Liao, M., G. Li, J. Wang, D. Marshall, T. Hui, S. Ma, Y. Zhang, B. Helmuth, and Y. Dong. 2021. Physiological determinants of biogeography: the importance of metabolic depression to heat tolerance. Global Change Biology, 27: 2561-2579. (Journal IF: 8.56).

Sara, G., M.C. Mangano, M. Berlino, L. Corbari, M. Lucchese, G. Millsenda, S. Terzo, M.S. Azaza, J.M.F. Babarro, R. Bakiu, B. Broitman, A.H. Buschmann, R. Christofoletti, A. Deidun, Y. Dong, J. Galdies, B. Glamuzina, O. Luthman, P. Makridis, A.J.A. Nogueira, M.G. Palomo, R. Dineshram, G. Rilov, P. Sanchez-Jerez, H. Sevgili, M. Troell, K.Y. AbouelFadl, M. Azra, P. Britz, C. Brugere, E. Carrington, I. Celic, F. Choi, Q. Chuanxin, T. Dobroslavic, P. Galli, D. Giannetto, J.H. Grabowski, M.J.H. Lebata-Ramos, P.T. Lim, Y. Liu, S.M. Llorens, G. Maricchiolo, S. Mirto, M. Pecarevic, N. Ragg, E. Ravagnan, D. Saidi, K. Schultz*, M. Shaltout, C. Solidoro, S.H. Tan, R.V. Thiyagarajan, and B. Helmuth. 2021. The synergistic impacts of anthropogenic stressors and COVID-19 on aquaculture: a current global perspective. Reviews in Fisheries Science and Aquaculture, doi: 10.1080/23308249.2021.1876633 (Journal IF: 3.95)


Helmuth, B., J.J. Leichter, R.D. Rotjan, K.D. Castillo, C. Fieseler, M.S. Jones and F. Choi. 2020. High resolution spatiotemporal patterns of seawater temperatures across the Belize Mesoamerican Barrier Reef. Nature Scientific Data, 7:396. doi: 10.1038/s41597-020-00733-6. (Journal IF: 4.80)

Mangano, M.C., N. Mieszkowska, B. Helmuth, T. Domingos, T. Sousa, G. Baiamonte, G. Bazan, A. Cuttitta, F. Fiorentino, A. Giacoletti, M. Johnson, G.D. Lucido, M. Marcelli, R. Martellucci, S. Mirto, B. Patti, F. Pranovi, G.A. Williams and G. Sarà. 2020. Moving towards a strategy for addressing climate displacement of marine resources: a proof-of-concept. Frontiers in Marine Science 7:408. doi: 10.3389/fmars.2020.00408 (Cited 4 times, Journal IF: 3.07)

Torossian, J.L., K.E. Hosek, S.C. Donelan, G.C. Trussell, B.S. Helmuth and M.L. Zippay. 2020. Physiological and biochemical responses to acute environmental stress and predation risk in the blue mussel, Mytilus edulis. Journal of Sea Research, 101891.

Wang, W., J. Wang, FMP Choi, P. Ding, X Li, G Han, M, Ding, M Guo, X. Huang, W. Duan, Z. Cheng, Z. Chen, S.J. Hawkins, Y. Jiang, B. Helmuth and Y. Dong. 2020. Global warming and artificial shorelines reshape seashore biogeography. Global Ecology and Biogeography 29 (2), 220-231.


Duwan, E., F. Choi and B. Helmuth. 2019. Assessing the Use of Virtual Reality Technology in Teaching Marine Ecological Concepts. Journal of STEM Outreach 2 (1), 1-11.

Speich et al. 2019. Global Observational Needs and Resources for Marine Biodiversity. Front. Mar. Sci. 6:367. doi: 10.3389/fmars.2019.00367

Choi, F., T. Gouhier, F. Lima, G. Rilov, R. Seabra and B. Helmuth. 2019. Mapping physiology: biophysical mechanisms define scales of climate change impacts. Conservation physiology 7(1).

Canonico et al. 2019. Global observational needs and resources for marine biodiversity. Frontiers in Marine Science. 6:367.

Rilov G., A.D. Mazaris, V. Stelzenmüller, B. Helmuth, M. Wahl, T. Guy-Haim, N. Mieszkowska, J. Ledoux and S. Katsanevakis. 2019. Adaptive marine conservation planning in the face of climate change: What can we learn from physiological, ecological and genetic studies? Global Ecology and Conservation 17 e00566.


Bates A.E., B. Helmuth, M.T. Burrows, M.I. Duncan, J. Garrabou, T. Guy-Haim, F. Lima, A.M. Queiros, R. Seabra, R. Marsh, J. Belmaker, N. Bensoussan, Y. Dong, A.D. Mazaris, D. Smale, M. Wahl and G. Rilov. 2018. Biologists ignore ocean weather at their peril. Nature 560(7718):299.

Connell S.D., Z.A. Doubleday, N.R. Foster, S.B. Hamlyn, C.D.G. Harley, B. Helmuth, B.P. Kelaher, I. Nagelkerken, K.L. Rodgers, G. Sarà and B.D. Russell. 2018. The duality of ocean acidification as a resource and a stressor. Ecology 99(5):1005-1010.

Judge R., F. Choi and B. Helmuth. 2018 Recent Advances in Data Logging for Intertidal Ecology. Front. Ecol. Evol. 6:213.

Zhou X., X. Ji, B. Wang, Y. Cheng, Z. Ma, F. Choi, B. Helmuth and W. Xu. 2018. Pido: Predictive Delay Optimization for Intertidal Wireless Sensor Networks. Sensors 18(5):1464.


Colvard N. and B. Helmuth. 2017. Nutrients influence the thermal ecophysiology of an intertidal macroalga: multiple stressors or multiple drivers? Ecological Applications 27(2): 669-681.

Connell S.D., Z.A. Doubleday, S.B. Hamlyn, N.R. Foster, C.D.G. Harley, B. Helmuth, B.P. Kelaher, I. Nagelkerken, G. Sarà and B.D. Russell. 2017. How ocean acidification can benefit calcifiers. Current Biology 27(3): R95-R96.

Dong Y., X. Li, F. Choi, G.A. Williams, G.N. Somero and B. Helmuth. 2017. Untangling the roles of microclimate, behaviour and physiological polymorphism in governing vulnerability of intertidal snails to heat stress. Proc. R. Soc. B 284(1854): 20162367.

Harley C.D.G., S.D. Connell, Z.A. Doubleday, B. Kelaher, B.D. Russell, G. Sarà and B. Helmuth. 2017. Conceptualizing ecosystem tipping points within a physiological framework. Ecology and Evolution DOI: 10.1002/ece3.3164.


Helmuth B., T.C. Gouhier, S. Scyphers and J. Mocarski. 2016. Trust, tribalism and tweets: has political polarization made science a “wedge issue”? Climate Change Responses 3(1): 3.

Helmuth B., F. Choi, A. Matzelle, J.L. Torossian,… 2016. Long-term, high frequency in situ measurements of intertidal mussel bed temperatures using biomimetic sensors. Scientific data 3: 160087.

Kish N.E., B. Helmuth and D.S. Wethey. 2016. Physiologically grounded metrics of model skill: a case study estimating heat stress in intertidal populations. Conservation physiology DOI: 10.1093/conphys/cow038.

Kroeker K.J., E. Sanford, J.M. Rose, C.A. Blanchette, F. Chan, F.P. Chavez, B. Gaylord, B. Helmuth, T.M. Hill, G.E. Hofmann, M.A. McManus, B.A. Menge, K.J. Nielsen, P.T. Raimondi, A.D. Russell and L. Washburn. 2016. Interacting environmental mosaics drive geographic variation in mussel performance and predation vulnerability. Ecology letters DOI: 10.1111/ele.12613.

Back to top



Monaco, C.J., D.S. Wetheyl, S. Gulledge and B. Helmuth. 2015. Shore-level size gradients and thermal refuge use in the predatory sea star Pisaster ochraceus: the role of environmental stressors. Marine Ecology Progress Series 539:191-205.

Marshall, D.J., E.L. Rezende, N. Baharuddin, F. Choi and B. Helmuth. 2015. Thermal tolerance and climate warming sensitivity in tropical snails. Ecology and Evolution 5(24):5905-5919.

Matzelle, A., G. Sará, V. Montalto, M. Zippay, G.C. Trussell and B. Helmuth. 2015. A Bioenergetics Framework for Integrating the Effects of Multiple Stressors: Opening a ‘Black Box’ in Climate Change Research. American Malacological Bulletin 33(1):150-160.

Back to top



Monaco, C.J., D.S. Wethey and B. Helmuth. 2014. A Dynamic Energy Budget (DEB) Model for the Keystone Predator Pisaster ochraceus. PLoS ONE 9(8): e104658. doi:10.1371/journal.pone.0104658.

Colvard, N.B., E. Carrington and B. Helmuth. 2014. Temperature-dependent photosynthesis in the intertidal alga Fucus gardneri and sensitivity to ongoing climate change . JEMBE 458:6-12.

Petes, L.E., J.F. Howard, B. Helmuth and E.K. Fly. 2014. Science integration into US climate and ocean policy. Nature Climate Change 4:671-677.

Montalto, V., G. Sará., P.M. Ruti, A. Dell’Aquila and B. Helmuth. 2014. Testing the effects of temporal data resolution on predictions of the effects of climate change on bivalves. Ecological Modelling 278:1-8.

Matzelle, A., V. Montalto, G. Sará, M. Zippay and B. Helmuth. 2014. Dynamic Energy Budget model parameter estimation for the bivalve Mytilus Californians: Application of the covariation method. Journal of Sea Research.

Sarà, G., M. Milanese, I. Prusina, A. Sarà, D.L. Angel, B. Glamuzina, T. Nitzan, S. Freeman, A. Rinaldi, V. Palmeri, V. Montalto, M.L. Martire, P. Gianguzza, V. Arizza, S.L. Brutto, M. De Pirro, B. Helmuth, J. Murray, S. De Cantis and G.A. Williams. 2014. The impact of climate change on Mediterranean intertidal communities: losses in coastal ecosystem integrity and services. Reg Environ Change 14:S5-S17.

Mislan, K.A., B. Helmuth and D.S. Wethey. 2014. Geographical variation in climatic sensitivity of intertidal mussel donation. Global Ecol. Biogeogr. 23:744-756.

Back to top



Sarà, G., V. Palmeri, A. Rinaldi, V. Montalto and B. Helmuth. 2013. Predicting biological invasions in marine habitats through eco‐physiological mechanistic models: a case study with the bivalve Brachidontes pharaonis. Diversity and Distributions 19(10):1235-1247.

Woodin, S.A., T.J. Hilbish, B. Helmuth, S.J. Jones and D.S. Wethey. 2013. Climate change, species distribution models, and physiological performance metrics: predicting when biogeographic models are likely to fail. Ecology and Evolution 3(10):3334-3346.

Pincebourde, S., E. Sanford and B. Helmuth. 2013. Survival and arm abscission are linked to regional heterothermy in an intertidal sea star. The Journal of Experimental Biology 216(12):2183-2191.

Ibáñez, I., E.S. Gornish, L. Buckley, D.M. Debinski, J. Hellmann, B. Helmuth, J. HilleRisLambers, A.M. Latimer, A.J. Miller-Rushing and M. Uriarte. 2013. Moving forward in global‐change ecology: capitalizing on natural variability. Ecology and Evolution 3(1):170-181.

Burnett, N. P., R. Seabra, M. d. Pirro, D. S. Wethey, S. A. Woodin, B. Helmuth, M. L. Zippay, G. Sarà, C. Monaco, and F. P. Lima. 2013. An improved noninvasive method for measuring heartbeat of intertidal animals. Limnology and Oceanography: Methods 11:91-100.

Back to top



Iacarella, J.C. and B. Helmuth. 2012. Body temperature and desiccation constrain the activity of Littoraria irrorata within the Spartina alterniflora canopy. Journal of Thermal Biology 37: 15-22.

Kearney, M., A. Matzelle and B. Helmuth. 2012. Biomechanics meets the ecological niche: the importance of temporal data resolution . J. Exp. Biol. 215: 922-933.

Pincebourde, S., E. Sanford, J. Casas, and B. Helmuth. 2012. Temporal coincidence of environmental stress modulates predation rates. Ecology Letters, 15: 680-688.

Zippay, M. and B. Helmuth 2012. Effects of temperature change on mussels, Mytilus (Linnaeus, 1798). Integrative Zoology 7: 312-327.

Back to top



Helmuth, B., L. Yamane, S. Lalwani, A. Matzelle, A. Tockstein, and N. Gao. 2011. Hidden signals of climate change in intertidal ecosystems: What (not) to expect when you are expecting. J. Exp. Mar. Biol. Ecol. 400: 191-199.

Iacarella, J.C. and B. Helmuth. 2011. Experiencing the salt marsh environment through the foot of Littoraria irrorata: Behavioral responses to thermal and desiccation stresses. J. Exp. Mar. Biol. Ecol. 409: 143-153.

Monaco, C.J. and B. Helmuth. 2011. Tipping Points, Thresholds and the Keystone Role of Physiology in Marine Climate Change Research. Adv. Mar. Biol. 60: 123-162.

Sará, G., M. Kearney and B. Helmuth. 2011. Combining heat-transfer and energy budget models to predict thermal stress in Mediterranean intertidal mussels. Chemistry and Ecology 27(2): 135-145.

Wethey, D.S., L.D. Brin, B. Helmuth and K.A.S. Mislan. 2011. Predicting intertidal organism temperatures with modified land surface models. Ecological Modelling 222: 3568-3576.

Zardi, G.I., K.R. Nicastro, C.D. McQuaid, L. Hancke and B. Helmuth. 2011. The combination of selection and dispersal helps explain genetic structure in intertidal mussels. Oecologia 165: 947-958.

Back to top



Helmuth, B., B.R. Broitman, L. Yamane, S.E. Gilman, K. Mach, K.A.S. Mislan, and M.W. Denny. 2010. Organismal climatology: analyzing environmental variability at scales relevant to physiological stress. J. Exp. Biol. 213: 995-1003.

Fuller, A., T. Dawson, B. Helmuth, R.S. Hetem, D. Mitchell and S. K. Maloney. 2010. Physiological mechanisms in coping with climate change. Physiol. Biochem. Zool. 83(5): 713-720.

Helmuth, B., L. Yamane, K.J. Mach, S. Chhotray, P. Levin and S. Woodin. 2010. All Climate Change is Local: Understanding and predicting the effects of climate change from an organism’s point of view. Stanford J. Law, Science, and Policy 2: 18-35.

Kearney, M., S.J. Simpson, D. Raubenheimer and B. Helmuth. 2010. Modelling the ecological niche from functional traits. Phil. Trans. Royal Society B 365: 3469-3483.

Schneider K.R., L.E. Van Thiel and B. Helmuth. 2010. Interactive effects of food availability and aerial body temperature on the survival of two intertidal Mytilus species. J. Thermal Biol. 35: 161-166.

Back to top



Broitman, B.R., P.L. Szathmary, K.A.S. Mislan, C.A. Blanchette and B. Helmuth. 2009. Predator-prey interactions under climate change: The importance of habitat vs body temperature. Oikos 118: 219-224.

Denny, M., and B. Helmuth. 2009. Confronting the physiological bottleneck: A challenge from ecomechanics. Integr. Comp. Biol. 49(3): 197-201.

Helmuth, B. 2009. From cells to coastlines: How can we use physiology to forecast the impacts of climate change? J. Exp. Biol. 212: 753-760.

Mislan, K.A.S., D.S. Wethey, and B. Helmuth. 2009. When to worry about the weather: Role of tidal cycle in determining patterns of risk in intertidal ecosystems. Global Change Biol. 15: 3056-3065.

Pincebourde, S., E. Sanford and B. Helmuth. 2009. An intertidal sea star adjusts thermal inertia to avoid extreme body temperatures. The American Naturalist 174(6): 890-897.

Szathmary, P.L., B. Helmuth and D. S. Wethey. 2009. Climate change in the rocky intertidal zone: Predicting and measuring the body temperature of a keystone predator. Mar. Ecol. Prog. Ser. 374: 43-56.

Yamane, L. and S.E. Gilman. 2009. Opposite responses by an intertidal predator to increasing aquatic and aerial temperatures. Mar. Ecol. Prog. Ser. 393: 27-36.

Back to top



Broitman, B.R., N. Mieszkowska, B. Helmuth and C.A. Blanchette. 2008. Climate and recruitment of rocky shore intertidal invertebrates in the Eastern North Atlantic. Ecol. 89(11) Supplement: S81-S90.

Pincebourde, S., E. Sanford and B. Helmuth. 2008. Body temperature during low tide alters the feeding performance of a top intertidal predator. Limnol. Oceanogr. 53(4): 1562-1573.

Back to top



Blanchette, C.A., S.D. Gaines and B. Helmuth. 2007. Spatial patterns of growth in the mussel, Mytilus Californians, across a major oceanographic and biogeographic boundary at Point Conception, California, USA. J. Exp. Mar. Biol. Ecol. 340: 126-148.

Finelli, C.M., B.S. Helmuth, N.D. Pentcheff and D.S. Wethey. 2007. Intracolony variability in photosynthesis by corals is affected by water flow: Role of oxygen flux. Mar. Ecol. Prog. Ser. 349: 103-110.

Helmuth, B. 2007. Forecasting the impacts of climate change on coastal ecosystems: How do we integrate science and policy? Southeastern Environmental Law Journal 16: 207-219.

Helmuth, B. 2007. Intertidal life as experienced through a powerful lens (Review of M. Koehl, “Wave-swept shore: The rigors of life on a rocky coast.”) Ecol. 88(1): 264-265.

Jost, J. and B. Helmuth. 2007. Morphological and ecological determinants of body temperature of Geukensia demissa, the Atlantic ribbed mussel, and their effects on mussel mortality. Biol. Bull. 213: 141-151.

Schneider, K.R. and B. Helmuth. 2007. Spatial variability in habitat temperature may drive patterns of selection between an invasive and native mussel species. Mar. Ecol. Prog. Ser. 339: 157-167.

Back to top



Finelli, C.M., B.S.T. Helmuth, N.D. Pentcheff and D.S. Wethey. 2006. Water flow influences oxygen transport and photosynthetic efficiency in corals. Coral Reefs 25: 47-57.

Gilman, S., C.D.G. Harley, D. Strickland, O. Vanderstraeten, M. O’Donnell and B. Helmuth. 2006. Evaluation of “Effective Shore Level” as a method of characterizing intertidal wave exposure regimes. Limnol. Oceanogr. Methods 4: 448-457.

Gilman, S.E., D.S. Wethey and B. Helmuth 2006. Variation in the sensitivity of organismal body temperature to climate change over local and geographic scales. Proc. Natl. Acad. Sci. 103: 9560-9565.

Helmuth, B., B.R. Broitman, C.A. Blanchette, S. Gilman, P. Halpin, C.D.G. Harley, M.J. O’Donnell, G.E. Hofmann, B. Menge and D. Strickland. 2006. Mosaic patterns of thermal stress in the rocky intertidal zone: Implications for climate change. Ecol. Monogr. 76: 461-479.

Helmuth, B., N. Mieszkowska, P. Moore and S.J. Hawkins. 2006. Living on the edge of two changing worlds: Forecasting the responses of rocky intertidal ecosystems to climate change. Ann. Rev. Ecol. Evol. Syst. 37: 373-404.

Leichter, J.J., B. Helmuth and A. Fischer. 2006. Variation beneath the surface: Quantifying complex thermal environments on coral reefs in the Caribbean, Bahamas, and Florida. J. Mar. Res. 64: 563-588.

Rotjan, R.D., J.L. Dimond, D.J. Thornhill, J.J. Leichter, B. Helmuth, D.W. Kemp and S.M. Lewis. 2006. Chronic fish grazing impedes coral recovery after bleaching. Coral Reefs 25: 361-368.

Back to top



Castillo, K.D. and B.S.T. Helmuth. 2005. Influence of thermal history on response of Montastraea annularis to short-term temperature exposure. Mar. Biol. 148: 261-270.

Helmuth, B., J.G. Kingsolver and E. Carrington, 2005. Biophysics, physiological ecology, and climate change: Does mechanism matter? Ann. Rev. Physiol. 67: 177-201.

Schneider, K. R., D. S. Wethey, B. Helmuth and T. J. Hilbish. 2005. Implications of movement behavior on mussel dislodgement: Exogenous selection in a Mytilus spp. hybrid zone. Mar. Biol. 146: 333-343.

Back to top



Denny, M.W. B. Helmuth, G.L. Leonard, C.D.G. Harley, L.Hunt and E. Nelson. 2004. Quantifying scale in ecology: Lessons from a wave-swept shore. Ecol. Monogr. 74: 513-532.

Fitzhenry, T., P.M. Halpin and B. Helmuth. 2004. Testing the effects of wave exposure, site, and behavior on intertidal mussel body temperatures: Applications and limits of temperature logger design. Mar. Biol. 145: 339-349.

Back to top



Denny, M.W., L.P. Miller, M.D. Stokes, L.J.H. Hunt and B.S.T. Helmuth. 2003. Extreme water velocities: Topographical amplification of wave-induced flow in the surf zone of rocky shores. Limnol. Oceanogr. 48: 1-8.

Harley, C.D.G. and B.S.T. Helmuth. 2003. Local and regional scale effects of wave exposure, thermal stress, and absolute vs. effective shore level on patterns of intertidal zonation. Limnol. Oceanogr. 48: 1498-1508

Helmuth, B. and M.W. Denny. 2003. Predicting wave exposure in the rocky intertidal zone: Do bigger waves always lead to larger forces? Limnol. Oceanogr. 48: 1338-1345.

Sebens, K.P., B. Helmuth, E. Carrington and B. Agius. 2003. Effects of water flow on growth and energetics of the scleractinian coral Agaricia tenuifolia, in Belize. Coral Reefs 22(1): 35-47.

Back to top



Helmuth B. 2002. How do we measure the environment? Linking intertidal thermal physiology and ecology through biophysics. Int. Comp. Biol. 42: 837-845.

Helmuth, B., C.D.G. Harley, P. Halpin, M. O’Donnell, G.E. Hofmann and C. Blanchette. 2002. Climate change and latitudinal patterns of intertidal thermal stress. Science 298: 1015-1017.

Tomanek, L. and B. Helmuth. 2002. Physiological ecology of rocky intertidal organisms: A synergy of concepts. Integ. Comp. Biol. 42: 771-775.

Back to top



Helmuth B. and G.E. Hofmann. 2001. Microhabitats, thermal heterogeneity and patterns of physiological stress in the rocky intertidal zone. Biol. Bull. 201: 374-384.

Back to top



Helmuth B. 1999. Thermal biology of rocky intertidal mussels: Quantifying body temperatures using climatological data. Ecol. 80: 15-34.

Back to top



Denny M.W., B. Gaylord, B. Helmuth and T.L. Daniel. 1998. The menace of momentum: Dynamic forces on flexible organisms. Limnol. Oceanogr. 43: 955-968.

Helmuth B.S.T. 1998. Intertidal mussel microclimates: Predicting the body temperature of a sessile invertebrate. Ecol. Monogr. 68: 29-52.

Sebens K.P., S.P. Grace, B. Helmuth, E.A. Maney, Jr. and J.S. Miles 1998. Water flow and prey capture by three scleractinian corals, Madracis mirabilis, Montastrea cavernosa and Porites porites in a field enclosure. Mar. Biol. 131: 347-360.

Back to top



Daniel T.L., B.S. Helmuth, W.B. Saunders and P.D. Ward. 1997. Septal complexity in ammonoid cephalopods increased mechanical risk and limited depth. Paleobiol. 23: 470-481.

Helmuth B.S.T., B.E.H. Timmerman and K.P. Sebens. 1997. Interplay of host morphology and symbiont microhabitat in coral aggregations. Mar. Biol. 130: 1-10.

Helmuth B.S.T., E.F. Stockwell and D.R. Brumbaugh. 1997. Morphological and environmental determinants of mass flux to corals. Proc. 8th Int. Coral Reef Symp., Panama 2: 1103-1108.

Helmuth B.S.T., K.P. Sebens and T.L. Daniel. 1997. Morphological variation in coral aggregations: Branch spacing and mass flux to coral tissues. J. Exp. Mar. Biol. Ecol. 209: 233-259.

Sebens K.P., J. Witting and B. Helmuth. 1997. Effects of water flow and branch spacing on particle capture by the reef coral Madracis mirabilis (Duchassaing and Michelotti). J. Exp. Mar. Biol. Ecol. 211: 1-28.

Back to top



Holberton R.L., B. Helmuth and J.C. Wingfield. 1996. The corticosterone stress response in Gentoo and King penguins during the non-fasting period. Condor 98: 850-854.

Padilla D.K., C.D. Harvell, J. Marks and B. Helmuth. 1996. Inducible aggression and intraspecific competition for space in a marine bryozoan, Membranipora membranacea. Limnol. Oceanogr. 41: 505-512.

Back to top



Helmuth B., R.R. Veit and R. Holberton. 1994. Dispersal of benthic invertebrates in the Scotia Arc by kelp rafting. Antarctic J. U.S. 29: 145-147.

Helmuth B., R.R. Veit and R. Holberton. 1994. Long-distance dispersal of a subantarctic brooding bivalve (Gaimardia trapesina) by kelp-rafting. Mar. Biol. 120: 421-426.

Back to top



Helmuth B. and K.P. Sebens. 1993. The influence of colony morphology and orientation to flow on particle capture by the scleractinian coral Agaricia agaricites (Linnaeus). J. Exp. Mar. Biol. Ecol. 165: 251-278.

Back to top